The family Charadriidae is very large with seventy species listed and is split into subfamilies.
The subfamily Charadriinae contains forty-two species in eight genera;
Charadrius (11), Anarhynchus (25), Oreopholus (1) Phegornis (1), Hoploxypterus (1) Eudromias (1), Zonibyx (1) and Peltohyas (1). We have further divided this group into three sections.
Section 1 Ringed Plovers.
Family: CHARADRIIDAE (Leach 1820)
Subfamily: CHARADRIINAE (Leach 1820)
Genera: There are eleven species of plover within the Charadrius genus.
Charadrius – Late Latin; charadrius yellowish bird mentioned in the Vulgate Bible (late 4th century) (Greek; kharadrios unknown plain-coloured nocturnal birds that dwelt in ravines and river valleys (kharadra ravine) (Linnaeus 1758)
- Europe: Common Ringed, Little Ringed.
- Africa: Common Ringed, Little Ringed, Three-banded, Forbes’.
- Asia: Common Ringed, Long-billed, Little Ringed.
- Australasia: Hooded, Shore, Black-fronted.
- North America: Common Ringed, Semipalmated, Killdeer, Piping.
- Central and South America: Semipalmated, Killdeer, Piping.
Genus Charadrius
∗ C. h. hiaticula (Linnaeus 1758)
∗ C. h. tundrae (Lowe 1915)
∗ C. h. psammodromus (Salomonsen 1930)
- Semipalmated Plover C. semipalmatus (Bonaparte 1825)
- Long-billed Plover C. placidus (Gray, J. & Gray, G. 1863)
- Little Ringed Plover C. dubius (Scopoli 1786)
∗ C. d. dubius (Scopoli 1786)
∗ C. d. curonicus (Gmelin 1789)
∗ C. d. jerdoni (Legge 1880)
- Killdeer C. vociferus (Linnaeus 1758)
∗ C. v. vociferus (Linnaeus 1758)
∗ C. v. ternominatus (Bangs & Kennard 1920)
∗ C. v. peruvianus (Chapman 1920)
- Piping Plover C. melodus (Ord 1824)
∗ C. m. melodus (Ord 1824)
∗ C. m. circumcinctus (Ridgway 1874)
∗ C. t. tricollaris (Vieillot 1818)
∗ C. t. bifrontalis (Cabanis 1882)
- Forbes’s Plover C. forbesi (Shelley 1883)
- Hooded Plover C. cucullatus (Vieillot 1818)
- Shore Plover C. novaeseelandiae (Gmelin 1789)
- Black-fronted Plover C. melanops (Vieillot 1818)
Genus Anarhynchus
- Wrybill Anarhynchus frontalis (Quoy & Gaimard 1830)
- Northern Red-breasted Plover Anarhynchus aquilonius (Gmelin 1789)
- Southern Red-breasted Plover A. obscurus (Dowding 1994)
- Wilson’s Plover A. wilsonia (Ord 1814)
∗ A. w. wilsonia (Ord 1814)
∗ A. w. beldingi (Ridgway 1919)
∗ A. w. cinnamominus (Ridgway 1919)
∗ A. w. crassirostris (Spix 1825)
- Madagascan Plover A. thoracicus (Richmond1896)
- Kittlitz’s Plover A. pecuarius (Temminck 1823)
- St Helena Plover A. sanctaehellenae (Harting 1873)
∗ A. m. marginatus (Vieillot 1818)
∗ A. m. mechowi (Cabanis 1884)
∗ A. m. arenaceus (Clancey 1971)
∗ A. m. tenellus (Hartlaub 1861)
∗ A. a. alexandrinus (Linnaeus 1758)
∗ A. a. seebohmi (Strickland 18152)
∗ A. a. nihonensis (Fischer & Reichenow 1884)
- Snowy Plover A. nivosus (Cassin 1858)
∗ A. n. nivosus (Cabanis 1872)
∗ A. n. occidentalis (Cabanis 1872)
- Javan Plover A. javanicus (Chasen 1938)
- White-faced Plover A. dealbatus (Swinhoe 1870)
- Red-capped Plover A. ruficapillus (Temminck 1822)
- Malaysian Plover A. peronii (Schlegel 1865)
- Chestnut-banded Plover A. pallidus (Strickland 1852)
∗ A. p. pallidus (Strickland 1852)
∗ A. p. venustus (Fischer & Reichenow 1884)
- Collared Plover A. collaris (Vieillot 1818)
- Puna Plover A. alticola (Berlepsh & Stolzmann 1902)
- Two-banded Plover A. falklandicus (Latham 1790)
- Double-banded Plover A. bicinctus (Jardine & Selby 1827)
∗ A. b. bicinctus (Jardine & Selby 1827)
∗ A. b. exilis (Falla 1978)
∗ A. m. mongolus (Pallas 1776)
∗ A. m. stegmanni (Portenko 1939)
∗ A. a. pamirensis (Richmond 1896)
∗ A. a. atrifrons (Wagler 1829)
∗ A. a. schaeferi (Meyer de Schauensee 1938)
∗ A. l. leschenaultii (Lesson 1826)
∗ A. l. columbinus (Wagler 1829)
∗ A. l. scythicus (Carlos, Roselaar & Voisin, 2012)
- Caspian Plover A. asiaticus (Pallas 1773)
- Oriental Plover A. veredus (Gould 1848)
- Mountain Plover A. montanus (Townsend 1837)
Ægialitis or Aegialitis (Boie 1822) is an old synonym for Charadrius but is no longer recognised in favour of the latter.
Wilson’s Plover subspecies Charadrius (now Anarhynchus). w. crassirostris (Carlos and Voisin 2011) was formerly called C. w. brasiliensis (Grantsau 2008). According with the International code of Zoological Nomenclature (1999) this change meant that Greater Sandplover subspecies C. l. crassirostris (Severtzov 1873) had to be remnamed C. l. scythicus (Carlos, Roselaar & Voisin 2012).
Mongolian and Lesser Sandplover were considered subspecies of Lesser Sandplover Charadrius mongolus. In 2022 a study showed that they were sufficiently different to warrant specific status creating Siberian Sandplover C. mongolus/stegmanniand and Tibetan Sandplover C. atrifrons/schaeferi/pamirensis. Both now in the genus Anarhynchus joining the only species originally in this genus Wrybill A. frontalis. Full List;
Northern Red-breasted Plover A. aquilonius
Southern Red-breasted Plover A. obscurus
Wilson’s Plover A. wilsonia
Madagascar Plover A. thoracicus
Kittlitz’s Plover A. pecuarius
St Helena Plover A. sanctaeheleniae
Kentish Plover A. alexandrinus
Snowy Plover A. nivosus
Javan Plover A. javanicus
White-faced Plover A. dealbatus
White-fronted Plover A. marginatus
Red-capped Plover A. ruficapillus
Malaysian Plover A. peronii
Chestnut-banded Plover A. pallidus
Collared Plover A. collaris
Puna Plover A. alticola
Northern and Southern Red-breasted Plover were formerly called New Zealand Dotterel and were considered subspecies Charadrius obscurus Obscurus and Charadrius o. aquilonius.
- Avibase https://avibase.bsc-eoc.org
- BirdLife International. IUCN Red List for birds. Downloaded from http://www.birdlife.org on 26/03/2020
- Carlos, C. J. & J-F. Voisin (2012). Charadrius wilsonia brasiliensis Grantsau & Lima, 2008, is a junior synonym of Charadrius crassirostris Spix, 1825: Bulletin of the British Ornithologists’ Club. 131 (1): 165–170.
- Carlos, C. J., S. Roselaar & J-F. Voisin (2012). A replacement name for Charadrius leschenaultii crassirostris (Severtzov, 1873), a subspecies of Greater Sand Plover: Bulletin of the British Ornithologists’ Club. 132 (1): 63–65.
- del Hoyo, Josep, Andrew Elliot & Jordi Sargatal. Handbook of the Birds of the World – Vol. 3 (1996)
- Grantsau, R. & P. C. Lima (2008). Uma nova subespécie de Charadrius wilsonia (Aves, Charadriiformes) para o Brasil: Atualidades Ornitológicas. 142: 4–5.
- Hayman, Peter, John Marchant & Tony Prater. Shorebirds – An identification guide to the waders of the world (1986)
- HBW Alive https://www.hbw.com
- Jobling, James A. Helm Dictionary of Scientific Bird Names (2010)
- Kennerley, P. R., D. N.Bakewell , P. D. Round. Rediscovery of a long-lost Charadrius plover from South-East Asia. Forktail 24, 63–79 (2008)
- Seebohm, Henry. The Geographical Distribution of the Family Charadriidae, or, The Plovers, Sandpipers, Snipes, and Their Allies (1888)
- Wei, Chentao, Manuel Schweizer, Pavel S Tomkovich, Vladimir Yu Arkhipov, Michael Romanov, Jonathan Martinez, Xin Lin, Naerhulan Halimubieke, Pinjia Que, Tong Mu, Qin Huang, Zhengwang Zhang, Tamás Székely, Yang Liu. Genome-wide data reveal paraphyly in the sand plover complex (Charadrius mongolus/leschenaultii) (2022)
Click on an image for more information about the individual species.
Genus Charadrius
Genus Anarhynchus